Gene Information
Gene NameUSP4
OrganismHomo sapiens (Human)
Gene Length963
Protein NamesUbiquitin carboxyl-terminal hydrolase 4 (EC 3.4.19.12) (Deubiquitinating enzyme 4) (Ubiquitin thioesterase 4) (Ubiquitin-specific-processing protease 4) (Ubiquitous nuclear protein homolog)
Target NameNA
Target TypeNA
Gene Age1714 ( ENSG00000114316 )
Evolutionary StageEukaryota
AlphaFoldDBQ13107
Gene CardUSP4
Uniprot IDQ13107
PfamPF06337; PF14836; PF00443
In ASDno
Allosteric Predictionyes (0.132527111)
Ortholog in AnimalsAlpaca_ USP4 ( ortholog_one2one )
Dog_ USP4 ( ortholog_one2one )
GuineaPig_ USP4 ( ortholog_one2one )
Macaque_ USP4 ( ortholog_one2one )
Mouse_ Usp4 ( ortholog_one2one )
Pig_ USP4 ( ortholog_one2one )
Rat_ Usp4 ( ortholog_one2one )
Rabbit_ USP4 ( ortholog_one2one )
OhnologsUSP11, USP15
ParalogsUSP11, USP20, USP5, USP42, USP14, USP29, USP25, USP48, USP2, USP47, USP40, USP3, USP1, USP34, USP8, USP21, USP24, USP38, USP32, USP6, USP31, USP50, USP13, USP7, USP43, USP17L4, USP17L11, USP17L27, USP17L22, USP17L30, USP17L17, USP17L26, USP17L25, USP17L12, USP17L10, USP17L28, USP17L1, USP17L5, USP17L24, USP17L21, USP17L3, USP17L29, USP17L18, USP17L13, USP17L15, USP17L7, USP17L23, USP17L2, USP17L20, USP17L8, USP17L19, USP35, USP45, USP33, USP49, USP51, USP19, USP15, USP28, USP9X, USP9Y, USP27X, USP44, USP18, USP30, USP10, USP22, USP26, USP16, USP37, USP36
TissueNA
Tissue SpecificityOverexpressed in small cell tumors and adenocarcinomas of the lung compared to wild-type lung (at protein level). Expressed in the hippocampal neurons. {ECO:0000269|PubMed:16339847, ECO:0000269|PubMed:7784062}.
Pharmacological Animal ModelsNA
Gene Ontology
(biological process)
Gene Ontology
(cellular component)
Gene Ontology
(molecular function)
PDB 2Y6E  (X-ray  2.40A  5CTR  (X-ray  3.01A 
DBSNP rs9311440 
Subcellular location [CC]Cytoplasm {ECO:0000269|PubMed:16316627, ECO:0000269|PubMed:20595234}. Nucleus {ECO:0000269|PubMed:16316627, ECO:0000269|PubMed:20595234}. Note=Shuttles between the nucleus and cytoplasm. Exported to the cytoplasm in a CRM1-dependent manner and recycled back to the nucleus via the importin alpha/beta heterodimeric import receptor. The relative amounts found in the nucleus and cytoplasm vary according to the cell type. {ECO:0000269|PubMed:16316627, ECO:0000269|PubMed:20595234}.
Mouse PG classificationNA
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