Gene Information
Gene NameSPAG9
OrganismHomo sapiens (Human)
Gene Length1321
Protein NamesC-Jun-amino-terminal kinase-interacting protein 4 (JIP-4) (JNK-interacting protein 4) (Cancer/testis antigen 89) (CT89) (Human lung cancer oncogene 6 protein) (HLC-6) (JNK-associated leucine-zipper protein) (JLP) (Mitogen-activated protein kinase 8-interacting protein 4) (Proliferation-inducing protein 6) (Protein highly expressed in testis) (PHET) (Sperm surface protein) (Sperm-associated antigen 9) (Sperm-specific protein) (Sunday driver 1)
Target NameNA
Target TypeNA
Gene Age987 ( ENSG00000008294 )
Evolutionary StageEumetazoa
AlphaFoldDBO60271
Gene CardSPAG9
Uniprot IDO60271
PfamPF16471; PF09744
In ASDno
Allosteric Predictionno (-0.411584347)
Ortholog in AnimalsAlpaca_ SPAG9 ( ortholog_one2one )
Dog_ SPAG9 ( ortholog_one2one )
GuineaPig_ SPAG9 ( ortholog_one2one )
Macaque_ SPAG9 ( ortholog_one2one )
Mouse_ Spag9 ( ortholog_one2one )
Pig_ SPAG9 ( ortholog_one2one )
Rat_ Spag9 ( ortholog_one2one )
Rabbit_ SPAG9 ( ortholog_one2one )
OhnologsMAPK8IP3
ParalogsMAPK8IP3
TissueNA
Tissue SpecificityExpressed only in testis on the round spermatids of stage I, II and II. Absent in spermatogonia and spermatocyte. {ECO:0000269|PubMed:9480848}.;
[Isoform 4]: Expressed in testis and in acute myeloid leukemia (AML) patients. {ECO:0000269|PubMed:16112646}.;
[Isoform 3]: Expressed in testis. {ECO:0000269|PubMed:14662895}.
Pharmacological Animal ModelsNA
Gene Ontology
(biological process)
Gene Ontology
(cellular component)
Gene Ontology
(molecular function)
PDB 2W83  (X-ray  1.93A 
DBSNP rs9896965 
Subcellular location [CC]Cytoplasm {ECO:0000250|UniProtKB:Q58A65}. Cytoplasm, perinuclear region {ECO:0000250|UniProtKB:Q58A65}. Lysosome membrane {ECO:0000269|PubMed:29146937}. Note=Perinuclear distribution in response to stress signals such as UV radiation. {ECO:0000250|UniProtKB:Q58A65}.;
[Isoform 5]: Cytoplasmic vesicle, secretory vesicle, acrosome {ECO:0000269|PubMed:15693750}. Note=Associated with the plasma membrane of the acrosomal compartment and also localizes in the acrosome matrix. {ECO:0000269|PubMed:15693750}.
Mouse PG classificationNA
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