Gene Information
Gene NameALYREF
OrganismHomo sapiens (Human)
Gene Length257
Protein NamesTHO complex subunit 4 (Tho4) (Ally of AML-1 and LEF-1) (Aly/REF export factor) (Transcriptional coactivator Aly/REF) (bZIP-enhancing factor BEF)
Target NameNA
Target TypeNA
Gene Age424 ( ENSG00000183684 )
Evolutionary StageEukaryota
AlphaFoldDBQ86V81
Gene CardALYREF
Uniprot IDQ86V81
PfamPF13865; PF00076
In ASDno
Allosteric Predictionno (-1.169352893)
Ortholog in AnimalsAlpaca_ ALYREF ( ortholog_one2one )
Dog_ ALYREF ( ortholog_one2one )
GuineaPig_ ALYREF ( ortholog_one2one )
Macaque_ ALYREF ( ortholog_one2one )
Mouse_ Gm4307 ( ortholog_one2many )
Mouse_ Alyref2 ( ortholog_one2many )
Mouse_ Gm8764 ( ortholog_one2many )
Mouse_ Gm6489 ( ortholog_one2many )
Mouse_ Alyref ( ortholog_one2many )
Mouse_ Gm21312 ( ortholog_one2many )
Mouse_ Gm4312 ( ortholog_one2many )
Mouse_ Gm4302 ( ortholog_one2many )
Mouse_ Gm4308 ( ortholog_one2many )
Mouse_ Gm20765 ( ortholog_one2many )
Mouse_ Gm21304 ( ortholog_one2many )
Mouse_ Gm4303 ( ortholog_one2many )
Mouse_ Gm4340 ( ortholog_one2many )
Mouse_ Gm21293 ( ortholog_one2many )
Mouse_ Gm4301 ( ortholog_one2many )
Mouse_ Gm4305 ( ortholog_one2many )
Mouse_ 4931428L18Rik ( ortholog_one2many )
Mouse_ Gm6763 ( ortholog_one2many )
Mouse_ Gm5698 ( ortholog_one2many )
Pig_ ALYREF ( ortholog_one2one )
Rat_ Alyref ( ortholog_one2one )
OhnologsNA
ParalogsPOLDIP3, CHTOP
TissueNA
Tissue SpecificityExpressed in a wide variety of cancer types. {ECO:0000269|PubMed:25662211}.
Pharmacological Animal ModelsNA
Gene Ontology
(biological process)
Gene Ontology
(cellular component)
Gene Ontology
(molecular function)
PDB 3ULH  (X-ray  2.54A 
DBSNP NA 
Subcellular location [CC]Nucleus {ECO:0000269|PubMed:10488337, ECO:0000269|PubMed:19324961, ECO:0000269|PubMed:23826332, ECO:0000269|PubMed:28418038, ECO:0000305|PubMed:18974867}. Nucleus speckle {ECO:0000269|PubMed:19324961, ECO:0000269|PubMed:23826332}. Cytoplasm {ECO:0000269|PubMed:19324961, ECO:0000269|PubMed:28418038}. Note=Colocalizes with the core EJC, ALYREF/THOC4, NXF1 and DDX39B in the nucleus and nuclear speckles. Travels to the cytoplasm as part of the exon junction complex (EJC) bound to mRNA (PubMed:19324961). Localizes to regions surrounding nuclear speckles known as perispeckles in which TREX complex assembly seems to occur (PubMed:23826332). {ECO:0000269|PubMed:19324961, ECO:0000269|PubMed:23826332}.
Mouse PG classificationNA
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